The "Floof-O-Meter" is basically a reputation system except you can't revoke your votes/take back your votes (You used to be able to and that's why you can see some people with negative numbers for the Floof-O-Meter). If you post content people like, such a helpful guides or just really cool stuff, people show you they like it by giving you a nuzzle, which brings the Floof-O-Meter up a point.