I think someone needs to start a new MMO, Aro'kai doesn't seem to be going anywhere, Feral-Heart seems to be at a standstill, Impressive-Title and its spinoffs are dominating that community, Aeon Phase isn't being worked on anymore, and WolfQuest.. is just eh.
I would say that the game I got most excited about was Aro'kai, until of course I waited all these years. Feral-Heart was the biggest surprise for me, I loved it upon release.
I don't think it's possible for me to compare these though... unless I discussed how things could be different.
I think we need more options in the animal-mmo community.