As well as how leader characteristics are applied to Feral Heart, I also think you can look to leaders outside of the game for similarities. Loyalty is a big one-- loyalty shows a bond between leader and groupmates, and I think repsect goes along with this, as well. I find that leaders who respect their pack/clanmates and leaders who understand that respect is earned rather than something they are entitled to is a very attractive quality, as far as leadership goes. Mutual respect and consideration for each individual goes a long way.
Humility is another quality that I find to be essential to a leader, in my eyes. While there may be some who disagree with this, as humility is often seen as weakness, I think it shows a trust and personal strength of the leader. Being humble is not always an easy thing to do, but it can be the difference between night and day, and possibly the difference between enjoying your time in a group and feeling as though you want to leave.
I think it's important for a leader to listen well-- actually listen, try to understand their groupmates' opinions and ideas before turning them away without a second thought, and have an open heart to the thoughts and feeling of their groupmates. It's one thing to listen to what someone says and forget it later, but if a leader is able to actually listen and seek to understand his groupmates, trust and respect can be drawn from this. I think this quality was already mentioned above, but this one is a biggie. I think communication falls into this category, as well. Keeping your group pals informed about whether you'll be absent a few days will speak volumes, whereas leaving your groupmates in the dark can cause unreliability and may make them feel as though you don't concern yourself with the wellbeing and interests of the pack.
Moral conscience-- While I think that it's important for a leader to listen to his group, having a firm grip on his own conscience and beliefs is also important. A leader who respects himself is more likely to be respected by others than one who does not have faith in his actions.
As a little side note, it's perfectly alright to remind yourself that you're there to have fun. Amidst all of the seriousness and pressure to try and "be" a good leader, at the end of the day, everyone is there to have fun and play around with creativity. It's never an "un-leaderly" thing to let go and just enjoy yourself.